Tag Archives: Tom Seeley

Bait hive location

We’re fast approaching the time of the season when colonies will attempt (and manage) to swarm. Swarming is the way bees increase their colony numbers.

Inevitably, because it means the loss of many of the foragers – the honey-gathering workers – beekeepers try and prevent or control swarming. Prevention is better than control, but it can be difficult or impossible to achieve.

Inevitably, a very large number of swarms are therefore lost.

The swarming process has been well-studied by eminent scientists like Martin Lindauer in the early 50’s and Tom Seeley more recently.

Swarming in a nutshell

The colony ‘decides’ to swarm. They rear one or more new queens in queen cells. On the 9th day after the egg is laid the cells are capped. Then or shortly afterwards the colony can be expected to swarm, with the original queen and up to half of the workers leaving with her. Swarming tends to happen on a warm, calm, sunny day (but not always). Casts may follow.

The swarm leaves the hive and settles nearby, in a tree or on a fencepost. Usually this initial ‘staging post’ is a short distance (perhaps 5-20 metres) from the hive. Scout bees fly from this swarm, surveying suitable locations in the area to set up a more permanent residence.

Swarm of bees

Swarm of bees

Actually, the scouts have been hard at work already. Once the swarm issues from the hive they set about persuading all the other scouts that they’ve found the ideal location. This ‘persuasion’ is the topic of the excellent book Honeybee Democracy by Tom Seeley.

Once a consensus is reached (which might take hours or a few days) the swarm leaves its temporary staging post and flies directly to the final location … a hollow tree, the church tower or above your neighbours soffit. There they set up home.

Most swarms do not survive. They get caught in a storm before relocating, they succumb to disease or they eventually starve to death. Since most strong colonies can – unchecked – be expected to swarm every year, the fact we’re not inundated with colonies indicates that most die.

One beekeepers loss can be another beekeepers gain.

Swarms can also be an annoyance to non-beekeepers. ‘Normal’ people. This always surprises me because a swarm is a stunning site and usually totally unthreatening and unaggressive 1.

Nevertheless, they can be an irritation, both when swarming or if they become established in the church tower or a hollow tree. And, without Varroa treatment they will act as a source of disease for other colonies in the locality through robbing and drifting 2.

So, rather than just let swarms disappear over the hedge or horizon, far better to capture them, put them in a hive and manage them properly 3.

There are essentially two ways to achieve this. Get them when they’re at their temporary staging post, drop them into a box and subsequently hive them. I’ve discussed this process before.

Captured swarm in 8 frame poly nuc

Captured swarm in 8 frame poly nuc …

Alternatively, set up one or more bait hives and entice a swarm to choose this location rather than the church tower or a hollow tree. Again, I’ve discussed how to set up a bait hives several times before.

Bait hives in a nutshell

Honeybee Democracy contains the definitive guide to the desirable characteristics of a bait hive. Conveniently it’s a dark box the size of a National brood box (~40 litres) with a small entrance (~12.5cm2) at the bottom. It helps a lot if it also smells of bees. This can easily be achieved by placing an old black brood frame up against one wall. Fill most or all of the rest of the box with foundationless frames 4 and place it in a suitable location.

Bait hive ...

Bait hive …

Which is – finally – what this post is all about.

Location, location, location

Seeley studied the desirable characteristics of both the bait hive and the location in which it is sited. By location I really mean height from the ground and orientation. However, he and others have also investigated how far from the original temporary staging post the swarm are prepared to fly to occupy a ‘des res’.

So, once you’ve prepared your bait hive(s), the three features of the location you need to think about are their a) orientation, b) height above ground, and c) distance from your bees (or other apiaries 5).

The first two are straightforward as Seeley describes in Honeybee Democracy. In the mid-70’s he conducted a series of experiments which showed that bees prefer bait hives that a) faced South and b) are located higher (5m from the ground) rather than lower (~1m) 6.

Most of the studies in Honeybee Democracy use artificially prepared swarms. Seeley would place these on a post and then observe where they relocated to … essentially he was circumventing the natural process of swarming by ‘starting’ them from the staging post. Many were conducted on an island where there were limited options to relocate to … and at limited distances.

These studies weren’t ideal to determine how far a swarm will relocate from its temporary staging post.

This is important in deciding whether to place bait hives in, near or distant from established hives.

Go forth and multiply, or vice versa

It might be logical to expect swarms to choose distant final destinations rather than nearby ones. For example, this might ensure less competition for forage. That would suggest that bait hives should be situated a long way from your own – or other – apiaries.

However, it turns out that when bees multiply they usually choose not to go forth very far. Although there is some contradictory evidence, it appears as though swarms prefer to relocate to nearby sites, rather than distant ones.

  1. Seeley and Morse (1977) demonstrated that swarms chose a nearby (~20m) site rather than a distant site (~400m) 80% of the time 7.
  2. Jaycox and Parise (1981) showed that swarms always chose sites less than 200m from the artificial swarms prepared for their study 8.
  3. Schmidt (1995) demonstrated that artificial swarms chose to relocate from as little as 0.5m to at least 1 km (average ~400m, i.e. much further away than the other studies) 9.

Why they prefer to set up home near to the original colony is unclear. At least one paper speculates it leaves the option to rob a genetically-related (and possibly less defensive) colony more easily.

Pragmatism and practical beekeeping

I set up bait hives every year. I’ve never failed to capture swarms. I’ve also – inadvertently – had swarms occupy stacks of stored brood boxes or supers. The entrances have faced almost every direction, with South East and South probably being the most often occupied.

However, this is a biased survey … I usually use the same site that worked in previous years.

Under offer ...

Under offer …

Although I’ve located bait hives high up on the top of greenhouses or sheds, the majority are on normal hive stands. This is pragmatism, not because they are more attractive to the bees at knee height 10.

I simply don’t want to climb a ladder to retrieve them 😉

Finally, I put them where I can see them and check them. I locate one in the corner of each apiary, as far from my other hives as possible, but probably never more than 10m away.

Pleasure and practical beekeeping

I also always have a bait hive in the garden where I can observe it on a daily basis. It’s great to see the first scouts appear, to watch their numbers increase day upon day, to keep an eye on the weather for a hot calm day … and to try and predict when the swarm will arrive.

Scout bees are not aggressive. They fly in a characteristic manner, checking the bait hive and the immediate vicinity from all directions.

If the numbers of scouts builds up to dozens or hundreds and they all disappear … either they’ve chosen somewhere else or the swam has been captured.

Or they’re guiding the swarm to your bait hive at that very moment.

Wait a few minutes. A swarm arriving is one of the really great sights in beekeeping 🙂


Bee lining for Christmas

Bee hunting

Bee hunting

Following the Wild Bees† by Tom Seeley is an entertaining little book that would make an ideal Christmas present for a beekeeper. It describes the methods used to locate feral colonies (or any colonies actually) by bee hunting or bee lining, so called because you follow the line or direction they return to the colony from a nectar source you provide. It’s an ideal Christmas book for two main reasons; it’s a summer activity, so will remind the reader that balmy sunny days will – finally – replace the cold, dark days of winter and, secondly, it will allow the enthusiast the time to build the essential two-chambered ‘bee lining box’ which is used to trap, feed and mark the bees being ‘lined’.

I don’t intend to provide a précis of the method … you should buy and read the book for that. However, as a taster, you can visit the companion website to the book or watch a short video of Tom Seeley bee hunting …

Tom Seeley is a Professor in the Department of Neurobiology and Behaviour at Cornell University. He is a highly respected entomologist and, unlike many scientists, writes in an engaging and accessible manner. He explains complicated experiments in layman’s terms and makes parallels between his observations on honey bees and wider societal issues. Anyone who has read his book “Honeybee Democracy” will appreciate how simple and elegant his description of the science is.

His explanation of bee hunting is no less clear. Following the Wild Bees is really a ‘how to’ guide, rather than a popular science book, though each chapter does contain a separate section on the science behind the ‘how to’, together with lots of anecdotes. The book is subtitled “The craft and science of bee hunting”. If you’re not aware of feral colonies in your own area this book might help you find them … however, if you live in an area with lots of other beekeepers it will probably just help you find their apiaries (and you can also do that with Google maps).

Wild? They’re livid feral.

The most up-to-date review of feral colonies in the UK can probably be found in Catherine Thompson’s 2012 doctoral thesis (brace yourself … this links to PDF of the 173 page thesis!). Catherine surveyed a number of feral colonies in the UK and showed that, although there were limited but significant genetic differences between feral colonies and managed colonies, the feral colonies were no more ‘native’. Catherine also neatly demonstrates the limitations of studying wing veination (morphometry) as an indicator of genetic purity – it usually isn’t. Feral colonies are essentially relatively recent swarms lost by local beekeepers.

Why ‘relatively recent’?

High levels of DWV

High levels of DWV …

The feral bees Catherine studied had much higher levels of deformed wing virus (DWV), both indicative of – and as would be expected of – uncontrolled Varroa infestation. Therefore, whilst it might appear appealing to have colonies of wild bees in the local church tower they’re almost certainly riddled with DWV and Varroa. This presumably explains why so many of the feral colonies Catherine analysed died during the study period (2.5 years). The swarms lost by beekeepers (that occupy the church tower for example) quickly succumb to the detrimental effects of uncontrolled Varroa replication and the consequent transmission of viruses. Furthermore, through the activities of robbing and drifting that feral colony is likely to act as the generous donor of viruses and mites to the local managed beekeepers hives.

Perhaps not so appealing after all.


I recommend you read Following the Wild Bees. Do so sitting in front of a roaring log fire in mid-winter. Plan and build a ‘bee lining box’ (or buy one) and consider where you might go prospecting for ‘wild’ bees once the long summer days return.

But also plan to put out bait hives to catch swarms (yours or others) and clip your queens … every one ‘lost’ is an opportunity to establish a future source of Varroa and virus infestation.

Under offer ...

Under offer …


 ISBN-10 0691170266 … it’s worth shopping around for a copy as the prices vary widely (at the time of writing). WH Smiths had it for well under a tenner recently.